■ Taxonomic and Host Catalogue of the Tachinidae of America North of Mexico
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Genus Carcelia Robineau-Desvoidy, 1830

CARCELIA Robineau-Desvoidy, 1830: 176. Type species: Carcelia bombylans Robineau-Desvoidy, 1830, by subsequent designation of Robineau-Desvoidy, 1863a: 220, 238 (as gnava Meigen, with bombylans in synonymy) [Palearctic].
GYMNOCARCELIA Townsend, 1919c: 582. Type species: Gymnocarcelia ricinorum Townsend, 1919 (= Eurigaster languida Walker, 1858), by original designation.
ELASSOMYIA Reinhard, 1967a: 103. Type species: Elassomyia defecta Reinhard, 1967 (= Eurigaster languida Walker, 1858), by original designation.


amplexa (Coquillett, 1897).– Southeastern Canada, eastern United States (Sabrosky & Reardon, 1976), Iowa (CNC).
– Exorista amplexa Coquillett, 1897: 98. Type data: holotype female (USNM). Type locality: USA, New Hampshire, White Mountains.
– Sisyropa hemerocampae Townsend, 1909: 248. Type data: holotype female (USNM). Type locality: USA, Massachusetts, Gypsy Moth Laboratory [at Melrose Highlands].

Hosts from Arnaud (1978: 114)*
Lepidoptera, Arctiidae
Ecpantheria eridanus (Cramer), Ecpantheria icasia (Cramer)
Lepidoptera, Lymantriidae
Euproctis chrysorrhoea (Linnaeus), Hemerocampa leucostigma (J.E. Smith), Orgyia antiqua (Linnaeus), Porthetria dispar (Linnaeus), Tussock moth(-s)
Lepidoptera, Megalopygidae
Megalopyge costaricensis Schaus, Megalopyge crispata (Packard), Megalopyge opercularis (J.E. Smith)
Lepidoptera, Noctuidae
Plathypena scabra (Fabricius), Spodoptera frugiperda (J.E. Smith)
Lepidoptera, Sphingidae
Manduca sexta jamaicensis (Butler)

diacrisiae Sellers, 1943.– Indiana to Connecticut, south to Alabama and Florida (S&A, 1965), Arizona, Texas (CNC).
– Carcelia diacrisiae Sellers, 1943: 61. Type data: holotype male (USNM). Type locality: USA, Indiana, Lafayette.

Hosts from Arnaud (1978: 115)*
Lepidoptera, Arctiidae
Diacrisia virginica (Fabricius), Estigmene acrea (Drury)

formosa (Aldrich & Webber, 1924).– Michigan to Massachusetts, south to Missouri and Florida, also Arizona, New Mexico (S&A, 1965), Illinois, Ontario, Québec (CNC).
– Zenillia (Zenillia) formosa Aldrich & Webber, 1924a: 23. Type data: holotype male (USNM). Type locality: USA, Massachusetts, West Medford.

Hosts from Arnaud (1978: 116)*
Lepidoptera, Noctuidae
Acronicta hamamelis (Guenée), Agrotis ipsilon (Hufnagel), Lithophane spp. (including L. disposita Morrison and/or L. innominata (Smith))
Lepidoptera, Saturniidae
Automeris io (Fabricius)

inflatipalpis (Aldrich & Webber, 1924).– British Columbia, Texas, Indiana, New York, Virginia, Florida (S&A, 1965), Missouri, Ontario, Québec (CNC).
– Zenillia (Zenillia) inflatipalpis Aldrich & Webber, 1924a: 24. Type data: holotype female (USNM). Type locality: USA, Virginia, Great Falls.


lagoae (Townsend, 1891).– Ohio, District of Columbia to Florida, also Texas, Mississippi, Mexico, Jamaica (S&A, 1965).
– Exorista lagoae Townsend, 1891c: 159. Type data: holotype male (SEMK). Type locality: Mexico, Guanajuato. Type host: Megalopyge opercularis (J.E. Smith) [published as Lagoa opercularis], Megalopygidae.

Hosts from Arnaud (1978: 117)*
Lepidoptera, Arctiidae
Ecpantheria icasia (Cramer), Halisidota spp.
Lepidoptera, Megalopygidae
Lagoa sp., Megalopyge crispata (Packard), Megalopyge krugii (Dewitz), Megalopyge opercularis (J.E. Smith), Megalopyge opercularis (J.E. Smith) probably
Lepidoptera, Pyralidae
Eulepte concordalis Hübner probably, Omphalocera cariosa Lederer

languida (Walker, 1858).– California to Ontario and New Hampshire, south to Mexico and Florida (S&A, 1965), Alberta to Manitoba (CNC).
– Tachina noctuae Harris, 1835: 599. Nomen nudum.
– Tachina albifrons Walker, 1853: 283 (junior homonym of Tachina albifrons Walker, 1836). Type data: holotype female (BMNH). Type locality: United States.
– Eurigaster languida Walker, 1858: 198 (genus cited as "Eurigaster ?"). Type data: holotype male (BMNH). Type locality: North America (as "Hindostan," in error; see Crosskey, 1976: 255).
– Gymnocarcelia ricinorum Townsend, 1919c: 582. Type data: lectotype male (USNM), by fixation of Townsend, 1941: 152 (mention of "Ht male" is regarded as a lectotype fixation). Type locality: USA, Florida, Allapata. Type host: Apantesis phyllira (Drury), Arctiidae.
– Elassomyia defecta Reinhard, 1967a: 103. Type data: holotype male (CNC). Type locality: USA, California.

Hosts from Arnaud (1978: 273), as Gymnocarcelia ricinorum Townsend*
Lepidoptera, Arctiidae
Apantesis phyllira (Drury), Diacrisia virginica (Fabricius), Ecpantheria deflorata (Fabricius), Estigmene acrea (Drury), Utetheisa ornatrix bella (Linnaeus)
Lepidoptera, Noctuidae
Pseudaletia unipuncta (Haworth)
Hosts from Arnaud (1978: 274) for probably languida Walker, as probably Gymnocarcelia ricinorum Townsend*
Lepidoptera, Arctiidae
Ecpantheria deflorata (Fabricius), Estigmene acrea (Drury)

laxifrons Villeneuve, 1912.– Palearctic; introduced and established in eastern North America (S&A, 1965; Richter & Wood, 1995), Québec (CNC).
– Carcelia laxifrons Villeneuve, 1912: 90, 91. Type data: holotype male (NHMW). Type locality: Europe (probably near Hamburg, Germany according to Herting, 1984: 187, note 42).

Hosts from Arnaud (1978: 118)*
Lepidoptera, Arctiidae
Apantesis proxima (Guérin-Méneville)
Lepidoptera, Lasiocampidae
Malacosoma americanum (Fabricius), Malacosoma disstria Hübner
Lepidoptera, Lymantriidae
Euproctis chrysorrhoea (Linnaeus), Nygmia phaeorrhoea (Donovan), Porthetria dispar (Linnaeus)
Lepidoptera, Notodontidae
Schizura concinna (J.E. Smith)

olenensis Sellers, 1943.– British Columbia, Québec, Massachusetts (S&A, 1965), California, Ontario, Newfoundland, Nova Scotia, North Carolina, Maine (CNC).
– Carcelia olenensis Sellers, 1943: 67. Type data: holotype male (USNM). Type locality: USA, Massachusetts, Saugus. Type host: Dasychira dorsipennata (Barnes & McDunnough) [published as Olene atomaria (Walker)], Lymantriidae.

Hosts from Arnaud (1978: 121)*
Lepidoptera, Lymantriidae
Dasychira atomeria Walker, Dasychira plagiata (Walker)

perplexa Sellers, 1943.– New York, Maine, Massachusetts, Connecticut, New Jersey (S&A, 1965).
– Carcelia perplexa Sellers, 1943: 65. Type data: holotype male (USNM). Type locality: USA, New Jersey, Trenton. Type host: Orgyia leucostigma (J.E. Smith) [published as Hemerocampa leucostigma], Lymantriidae.

Hosts from Arnaud (1978: 121)*
Lepidoptera, Lymnatriidae
Dasychira plagiata (Walker), Hemerocampa leucostigma (J.E. Smith)

protuberans (Aldrich & Webber, 1924).– Vermont (S&A, 1965), British Columbia to New Brunswick and New Jersey, also California, Colorado, Texas, Tennessee, Georgia (S&A, 1965, as malacosomae), Michigan, Ohio, New York (CNC).
– Zenillia (Zenillia) protuberans Aldrich & Webber, 1924a: 15. Type data: holotype male (USNM). Type locality: USA, Vermont, Rockingham. Type host: Malacosoma disstria Hübner, Lasiocampidae.
– Carcelia malacosomae Sellers, 1943: 50 (named for Exorista cheloniae of authors, not Rondani, 1859). Type data: holotype male (USNM). Type locality: USA, Massachusetts, Saugus.

Hosts from Arnaud (1978: 121)*
Lepidoptera, Arctiidae
Hyphantria cunea (Drury)
Lepidoptera, Lasiocampidae
Malacosoma americanum (Fabricius), Malacosoma disstria Hübner
Hosts from Arnaud (1978: 119), as Carcelia malacosomae Sellers*
Unidentified Hymenoptera
Unidentified Tenthredinidae
Lepidoptera, Arctiidae
Hyphantria cunea (Drury), Turuptiana permaculata (Packard)
Lepidoptera, Lasiocampidae
Malacosoma americanum (Fabricius), Malacosoma disstria Hübner, Datana integerrima Grote & Robinson
Lepidoptera, Saturniidae
Hemileuca spp. (including H. lucina H. Edwards and/or H. maia (Drury))
Hosts from Arnaud (1978: 120) for probably protuberans (Aldrich & Webber), as probably Carcelia malacosomae Sellers*
Lepidoptera, Arctiidae
Apantesis ornata (Packard), Apantesis proxima (Guérin-Méneville), Arachnis picta Packard, Turuptiana permaculata (Packard)
Lepidoptera, Lasiocampidae
Malacosoma americanum (Fabricius), Malacosoma disstria Hübner, American tent caterpillar(-s)
Lepidoptera, Tortricidae
Archips argyrospilus (Walker)

reclinata (Aldrich & Webber, 1924).– British Columbia, Ontario, all United States (S&A, 1965), Yukon, Northwest Territories, Alberta and Nunavut to New Brunswick (CNC).
– Zenillia (Parexorista) reclinata Aldrich & Webber, 1924a: 32. Type data: holotype male (USNM). Type locality: USA, New Mexico, Hell Canyon.

Hosts from Arnaud (1978: 122)*
Lepidoptera, Arctiidae
Diacrisia virginica (Fabricius), Ecpantheria deflorata (Fabricius), Estigmene acrea (Drury), Phragmatobia assimilans Walker
Hosts from Arnaud (1978: 123) for probably reclinata (Aldrich & Webber)*
Lepidoptera, Arctiidae
Estigmene acrea (Drury), Phragmatobia assimilans Walker

tenuiforceps (Reinhard, 1964).– Ohio (Reinhard, 1964), Ontario, Québec, Nova Scotia (CNC).
– Gymnocarcelia tenuiforceps Reinhard, 1964b: 49. Type data: holotype male (CNC). Type locality: USA, Ohio, Amherst.


yalensis Sellers, 1943.– British Columbia, Idaho, California, Ontario to Maine and Massachusetts (S&A, 1965), Yukon, Saskatchewan, Texas (CNC).
– Carcelia yalensis Sellers, 1943: 69. Type data: holotype female (USNM). Type locality: USA, Idaho, Yale.

Hosts from Arnaud (1978: 123)*
Lepidoptera, Arctiidae
Halisidota tessellaris (J.E. Smith)
Lepidoptera, Lymnatriidae
Hemerocampa leucostigma (J.E. Smith), Orgyia antiqua (Linnaeus)

Carcelia species

Hosts from Arnaud (1978: 124) for Carcelia spp.*
Lepidoptera, Limacodidae
Adoneta spinuloides Herrich-Schaeffer, Phobetron pithecium (J.E. Smith)
Lepidoptera, Noctuidae
Alypia octomaculata (Fabricius), Peridroma saucia (Hübner)
Lepidoptera, Notodontidae
Datana integerrima Grote & Robinson
Lepidoptera, Olethreutidae
Melissopus latiferreanus (Walsingham)
Lepidoptera, Pyralidae
Pantographa limata Grote & Robinson
Lepidoptera, Sphingidae
Manduca sexta (Linnaeus)

Reference:


Arnaud, P.H., Jr. 1978. A host-parasite catalog of North American Tachinidae (Diptera). United States Department of Agriculture. Miscellaneous Publication 1319: 1–860.


*Host names (family and species) have not been changed from those given in Arnaud (1978). Each host is listed under the appropriate current tachinid name, with the tachinid name used in Arnaud (1978) cited if different from the current one. For more information about Arnaud (1978), and to see a complete list of tachinid names used in that work and their modern equivalents, click here.

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